Whomever guessed paint in the axle sockets was apparently correct.
I had to dismantle the center truck, again (I’m getting quite adept at reassembly, disassembly happens more or less automatically once you back out the one tiny Phillips head screw), to repaired an axle retaining clip. The one pin snapped so I tapped the hole, extending thechole and the thresding through the plastic clip, and inserted an 00-90 brass machine screw from Hob-Its, clipping off the head and filing the stub flush.
Anyway, while it was apart I remembered someone suggesting paint in the axle holes and that turned out to be correct. Also on some axle stub ends which was easier to clean off. Reassembled with conductalube now accurately placed right in the axle holes and this Shay runs like new again.